Amplifying force.

Palm cockatoo (Probosciger aterrimus) leaf clamping

Wild tuna (Thunnus spp.) shark scraping

Long-tailed macaque (Macaca fascicularis aurea) crab hunting

Blanket octopus (Tremoctopus sp.) medusa carrying

Yellow-legged gull (Larus michahellis) clam dropping

Dinosaur gastroliths

Capuchin monkey (Sapajus libidinosus) stone flaking

Sea otter (Enhydra lutris) mussel pounding

Banded mongoose (Mungos mungo) millipede throwing

Warty pig (Sus cebifrons) nest digging

King Kong (Titanus kong) tool use

Captive orangutan (Pongo pygmaeus) stone flaking

Palm cockatoo (Probosciger aterrimus) stick drumming

Long-tailed macaque (Macaca fascicularis) nut cracking

Snapping shrimp (Synalpheus regalis) bubble stunning

Bearded capuchin (Sapajus libidinosus) stone digging

Robotic spacecraft (Hayabusa2) asteroid sampling

Graphic tuskfish (Choerodon graphicus) anvil striking

A RARE example

Defining tool use

Egyptian vulture (Neophron percnopterus) stone throwing